B.S., University of Wisconsin-Milwaukee, 1965.
M.S., University of Wisconsin-Milwaukee, 1967.
Ph.D., Michigan State University, 1972.

Hepatic systematics and phytogeography of the south temperate and subantarctic regions.

The primary objective of my research program is to come to an understanding of hepatic systematics and phytogeography of south temperate and subantarctic regions. These geographic areas present an ideal natural laboratory to investigate biological questions relevant to evolutionary persistence, dispersibility and survival and evolution of plants faced with environmental change. Within that framework I am engaged in detailed, in-depth (monographic) studies of various groups, including Chiloscyphus, Heterorcyphus and Telaranea for Australasia. My research also focuses on the “Hepaticae of the New Zealand”, a three-volume work in collaboration with David Glenny of Landcare, Lincoln, New Zealand. My program includes elements of biogeography, ecology, morphology and phylogeny. I am also investigating problems involving functional morphology of hepatic structures and I am doing so in a broad evolutionary context.