M.A., Sociology, University of Chicago, 1951.
M.Ed., Science Education, Chicago Teachers College, 1956.
Ph.D., Zoology, University of Chicago, 1960.

Consulting Editor, American Journal of Primatology.
Editorial Board, International Journal of Primatology.

Primate systematics, particularly Asian catarrhines.

My major research effort has been devoted to systematic review of the genus Macaca, a widely distributed group of Old World monkeys. Subjects investigated include geographic variation of external, cranial, and molecular characters; natural history; and paleontology. Field work has been conducted in collaboration with local zoologists in China, India, Indonesia and Thailand. To date, fifteen of the nineteen recognized species of macaques have been monographed.