B.S., Biology, University of Minnesota, 1971.
M.A., Systematics and Ecology, University of Kansas, 1975.
Ph.D., Systematics and Ecology, University of Kansas, 1979.

Research Fellow, Smithsonian Institution, 1986-1988.
Research Associate, Smithsonian Institution, 1988-present.
Research Associate, American Museum of Natural History, 1991-present.
Research Associate, Utah Museum of Natural History, 1994-present.
Honorary Curator, Department of Zoology, Philippine National Museum, 1990-present.
Science Advisor, Center for Tropical Conservation Studies, Silliman University, Philippines, 1992-present.

Mammalian evolution and ecology/evolutionary biogeography/origin and maintenance of patterns of biological diversity/conservation biology/tropical biology.

In spite of the long-term interest of biologists in the dynamics of the processes that influence patterns of biological diversity, a comprehensive understanding has yet to emerge. My current research program focuses on the ecology and evolution of mammals on the islands of Southeast Asia, especially the Philippines, as a natural laboratory where the influence of island area, habitat diversity, degree of isolation, and geological histories may be investigated. Variation in these factors allows direct investigation of patterns of extinction, colonization, and speciation in a remarkably diverse fauna. Studies include analysis of morphological, genetic, and ecological diversification, elevational diversity patterns, evolution of life history traits, reproductive ecology, and plant-animal interactions. Rapid destruction of natural habitats and extinction of native species has led me to integrate formal ecological and evolutionary studies with research and consulting activites on the impact of deforestation. I am actively involved in providing advanced training in conservation biology to scientists from the tropics.

Much of my work includes participation by graduate students from both within and outside of the United States. Past students have studied reproductive ecology of bats, seed dispersal systems, tree squirrel population biology, mammalian paleoecology, evolution and functional morphology of horned mammals, population genetics of mammals in island ecosystems and population biology of tropical birds.