B.S., Zoology, Iowa State University, 1970.
M.A., Systematics and Ecology, University of Kansas, 1976.
Ph.D., Systematics and Ecology, University of Kansas, 1979.

Fellow, American Ornithologists' Union; Program Coordinator (Assistant to Secretary of AOU).
Editor, Inland Bird Banding Association (for North American Bird Bander).

Brood parasitism and avian population biology

Research interests include study of breeding biology of birds at population and community levels and work to understand the ecology and evolution of avian brood parasitism and resultant interactions between host and parasite populations. Active projects include monitoring breeding biology of a suburban nesting colony of House Sparrows (since 1988) and study of brood parasitism by Brown-headed Cowbirds.

Checking nestboxes at suburban colony of House Sparrows