A.B., Chemistry, Rutgers University, 1966.
A.M., Chemistry, Harvard University, 1970.
Ph.D., Zoology, Harvard University, 1973.

Editorial Board, Psyche, 1978-1986.
Australian Biological Resources Study.
Councilor, Coleopterists Society (1993-1994).
Field Research program supported by NSF, American Philosophical Society, Ernst Mayr Grant, National Geographic Society.

Phylogeny, comparative morphology and evolution of beetles/historical biogeography/insect association with specialized microhabitats.

My current research revolves around studies on the evolution of the large beetle family Staphylinidae (over 45,000 named species). A long-term study of the higher-level classification and evolution of the group continues by focusing on reconstructing the phylogeny of one of the family's four main lineages. I have nearly completed a monographic study of the 190+ New World species of the genus Platydracus, whose species promise to be of special interest for understanding the origin of current distribution patterns of forest-dwelling insects in Mexico and Central America. Collaboration with M. K. Thayer continues on another long-term project to improve knowledge of the staphylinoid beetle fauna of Australia and other southern temperate areas, with the ultimate aim of using this group to help understand the origin of southern disjunct distribution patterns (Australia+New Zealand+Chile) that are very common within the group.