Ph.D., University of Chicago, 1967.

Bibliographic compiler for Saugetierkundliche Mitteilungen from 1955-65.
Vice-president and President of the Society of Vertebrate Paleontology, 1976-1977, 1977-1978.
Co-leader, three expeditions to Australia (1963-4), (1966-7), (1976-7) and leader of many field trips into the Washakie Basin of Wyoming (1956-96).

Mammalian systematics, evolution, ecology and zoogeography

Mammalian systematics, evolution, ecology and zoogeography, especially North American Eocene and Mesozoic faunas, Australian Tertiary and Pleistocene faunas/functional morphology of mammalian masticatory apparatus/paleopathology/taphonomy/basicranial anatomy of marsupials/group origins and replacements.

Current research involves: i) the mammalian fauna of the Washakie Formation (southwest Wyoming) and the stratigraphic sequence of the deposits of the formation (uintatheres, marsupials, titanotheres, artiodactyls, tillodonts, and several other works in progress); ii) Late Tertiary and Pleistocene mammalian faunae of Australia; iii) functional anatomy and restoration of the masticatory musculature of the Eocene taeniodont, Stylinodon,; and iv) the multituberculates of the Early Cretaceous (Trinity Group) of north Texas.